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He was the son of [[Sidney Coupland]], a physician at [[Middlesex Hospital]], and his wife Bessie Potter, daughter of Thomas Potter of [[Great Bedwin]], born in London. He was educated at [[Winchester College]], and went on [[New College, Oxford]], where he was taught by [[Alfred Zimmern]], among others. He graduated in 1907, with a first class in [[Greats]]. That year he was elected a Fellow at [[Trinity College, Oxford|Trinity College]] where he lectured in [[ancient history]].<ref name="ODNB"/>
Under the influence of [[Lionel Curtis]], Beit lecturer in colonial history 1912–1913, Coupland joined the [[Round Table movement]], and succeeded Curtis as Beit lecturer. He became Beit Professor in 1920, despite a lack of finished work in print.<ref name="ODNB"/>
With [[Lionel Curtis]], Coupland tried to set up an African institution in [[Rhodes House]] in the early 1930s; but they were unsuccessful in getting funding.<ref>{{cite book |last1=Hodge |first1=Joseph Morgan |title=Triumph of the Expert: Agrarian Doctrines of Development and the Legacies of British Colonialism |date=2007 |publisher=Ohio University Press |isbn=978-0-8214-4226-5 |page=138 |url=https://books.google.co.uk/books?id=gcWeBAAAQBAJ&pg=PA138 |language=en}}</ref> From 1938 to 1943 Coupland assisted [[Lord Lugard]] and Hanns Vischer with the running of the [[International African Institute]].<ref>''Sir Reginald Coupland, K.C.M.G., C.I.E.'', Africa: Journal of the International African Institute, Vol. 23, No. 1 (Jan., 1953), p. 1. Published by: Cambridge University Press on behalf of the International African Institute {{jstor|1156026}}</ref>▼
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Coupland took part in the [[Cripps Mission]] of 1942 to Indian leaders. His diary of 1941–1942 is a significant source for the activities and thinking of [[Sir Stafford Cripps]]. It also discusses the Indian political groups.<ref>{{cite book |last1=Jafri |first1=Saiyid Zaheer Husain |title=Recording the Progress of Indian History: Symposia Papers of the Indian History Congress, 1992-2010 |date=2012 |publisher=Primus Books |isbn=978-93-80607-28-3 |page=510 |url=https://books.google.co.uk/books?id=K4SzQL1uds4C&pg=PA510 |language=en}}</ref> He was closely involved with [[Graham Spry]] in contradicting the account published by [[Louis Fischer]] in ''[[The Nation]]'' of political undertakings given by Cripps to [[Abul Kalam Azad]], [[Mahatma Gandhi]] and [[Jawaharlal Nehru]].<ref>{{cite book |last1=Clarke |first1=Peter |title=The Cripps Version: The Life of Sir Stafford Cripps, 1889-1952 |date=2003 |publisher=Penguin Books, Limited |isbn=978-0-14-028691-5 |pages=352-353 |url=https://books.google.co.uk/books?id=NkE8AgAACAAJ&pg=PA352 |language=en}}</ref>
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