Cucurbitaceae

Cucurbitaceae; Sakup ti panawen: Nasapa a Paleoseno aginggana iti Kaudian, 62–0 Ma PreЄ Є O S D C P T J K Pg N
	Lalaki a mula a Hodgsonia
Taksonomia
Pagarian:	Plantae
Klado:	Tracheophytes
Klado:	Angiospermae
Klado:	Eudicotidae
Klado:	Rosids
Urnos:	Cucurbitales
Pamilia:	Cucurbitaceae; Juss.
Henero-kita
	Cucurbita; L.
Dagiti tribu ken henero
	Kitaen ti teksto.

Ti Cucurbitaceae matawtawagan pay iti cucurbits ken ti pamilia ti tabungaw, ket dagiti pamilia ti mula a mangbukel kadagiti agarup a 965 a sebbangan kadagiti agarup a 95 a henero,^[2] dagiti kangrunaan ket ti:

Cucurbita – karabasa, pumpkin, zucchini, ken dagiti dadduma a tabungaw
Lagenaria – calabash, ken dagiti dadduma a saan a makan
Citrullus – sandia (C. lanatus, C. colocynthis) ken dagiti dadduma
Cucumis – pipino (C. sativus), dagiti nadumaduma a melon
Luffa – luffa pay ti kadawyan a nagan, sagpaminsan a mailetra iti loofah

Dagiti mula iti daytoy a pamilia ket naimuyongan kadagiti tropiko ken kadagiti temperado a lugar, a dagiti addaan iti makan a bunga ket isuda ti kasapaan a naimulmula kadagiti Daan ken Baro a Lubong. Mairanggo ti pamilia ti Cucurbitaceae kadagiti kaaduan ti bilang a pamilia ti mula ken porsiento dagiti sebbangan nga inus-usar a makan iti tao.^[3]

Buklen ti Cucurbitaceae dagiti 98 a naisingasing a henero nga agraman kadagiti 975 a sebbangan,^[4] a naipangpangruna kadagiti rehion ti tropikal ken subtropikal. Amin dagiti sebbangan ket senitibo iti linaaw a nagbalay. Kaaduan kadagiti mula iti daytoy a pamilia ket dagiti tinawen nga agkalatkat, ngem adda dagiti natarikayo a liana, nasiit a bassit a kayo, wenno kaykayo (Dendrosicyos). Adu kadagiti sebbangan ket dakkel, duyaw wenno puraw a sabsabong. Nadutdotan ken pentangular dagiti ungkay. Adda dagiti pakalatkat iti 90° kadagiti pesiolo ti bulong kadagiti nodo. Dagiti bulong ket agsisinublat nga estipula a simple a palmato wenno dua palmato. Dagiti sabong ket uniseksual, nga addaan kadagiti lalaki ken babai a sabong kadagiti sabali a mula (dioecious) wenno iti isu met laeng a mula (monoecious). Dagiti babai a sabong ket inferior nga iitlogan. Ti bunga ket masansana kas ti nabaliwan a berry a tinawtawagan iti pepo.

Pakasaritaan ti posil

Maysa kadagiti kaduogan a posil ti cucurbits ket ti †Cucurbitaciphyllum lobatum manipud iti panawen ti Paleoseno, a nabirukan idiay Shirley Canal, Montana. Immuna a naipalawag daytoy idi 1924 babaen ti paleobotaniko a ni Frank Hall Knowlton. Ti posil a bulong ket palmato, tallo-rutong nga agraman kadagiti nagtimbukel a rutong ti sinus ken iti intero wenno rimmagadi a paraigid. Daytoy ket addaan iti sukog ti bulong a kapada dagiti kameng dagiti henero ti Kedrostis, Melothria ken Zehneria.^[5]

Pannakaidasig

Maipapan iti tribu a pannakaidasig

Ti kaadian a pannakaidasig ti Cucurbitaceae ket nalawag a mangipuesto kadagiti 15 a tribu:^[6]^[7]

Tribu ti Gomphogyneae Benth. & Hook. f. 1867
- Alsomitra (Blume) Spach 1838 (1 sp.)
- Bayabusua (1 sp.)
- Gomphogyne Griff. 1845 (2 spp.)
- Gynostemma Blume 1825 (10 spp.)
- Hemsleya Cogn. ex F.B. Forbes & Hemsl. 1888 (30 spp.)
- Neoalsomitra Hutch. 1942 (12 spp.)
Tribu ti Triceratieae A. Rich. 1845
- Anisosperma Silva Manso 1836 (1 sp.)
- Cyclantheropsis Harms 1896 (3 spp.)
- Fevillea L. 1753 (8 spp.)
- Pteropepon (Cogn.) Cogn. 1916 (5 spp.)
- Sicydium Schltdl. 1832 (7 spp.)
Tribu ti Zanonieae Benth. & Hook. f. 1867
- Gerrardanthus Harvey in Hook. f. 1867 (3–5 spp.)
- Siolmatra Baill. 1885 (1 sp.)
- Xerosicyos Humbert 1939 (5 spp.)
- Zanonia L. 1753 (1 sp.)

Tribu ti Actinostemmateae H. Schaef. & S.S. Renner 2011
- Actinostemma Griff. 1841 (3 spp.)

Tribu ti Indofevilleeae H. Schaef. & S.S. Renner 2011
- Indofevillea Chatterjee 1946 (1 sp.)
Tribu ti Thladiantheae H. Schaef. & S.S. Renner 2011
- Baijiania A.M. Lu & J.Q. Li 1993 (30 spp.)
- Thladiantha Bunge 1833 (5 spp.)
Tribu ti Siraitieae H. Schaef. & S.S. Renner 2011
- Siraitia Merr. 1934 (3–4 spp.)
Tribu ti Momordiceae H. Schaef. & S.S. Renner 2011
- Momordica L. 1753 (60 spp.)
Tribu ti Joliffieae Schrad. 1838
- Ampelosicyos Thouars 1808 (5 spp.)
- Cogniauxia Baill. 1844 (2 spp.)
- Telfairia Hook. 1827 (3 spp.)

Tribu ti Bryonieae Dumort. 1827
- Austrobryonia H. Schaef. 2008 (4 spp.)
- Bryonia L. 1753 (10 spp.)
- Ecballium A. Rich. 1824 (1 sp.)
Tribu ti Schizopeponeae C. Jeffrey 1964
- Herpetospermum Wall. ex Hook. f. 1867 (3 spp.)
- Schizopepon Maxim. 1859 (6–8 spp.)
Tribu ti Sicyoeae Schrad. 1838
- Cyclanthera Schrad. 1831 (40 spp.)
- Echinocystis Torr. & A. Gray 1840 (1 sp.)
- Echinopepon Naudin 1866 (20 spp., including Brandegea Cogn. 1890)
- Frantzia Pittier 1910 (5 spp.)
- Hanburia Seem. 1858 (7 spp.)
- Hodgsonia Hook. f. & Thomson 1854 (2 spp.)
- Linnaeosicyos H. Schaef. & Kocyan 2008 (1 sp.)
- Luffa Mill. 1754 (5–7 spp.)
- Marah Kellogg 1854 (7 spp.)
- Nothoalsomitra Hutch. 1942 (1 sp.)
- Sicyos L. 1753 (75 spp., including Sechium P. Browne 1756)
- Trichosanthes L. 1753 (≤100 spp.)

Tribu ti Coniandreae Endl. 1839
- Apodanthera Arn. 1841 (16 spp.)
- Bambekea Cogn. 1916 (1 sp.)
- Ceratosanthes Adans. 1763 (4 spp.)
- Corallocarpus Welw. ex Benth. & Hook. f. 1867 (17 spp.)
- Cucurbitella Walp. 1846 (1 sp.)
- Dendrosicyos Balf. f. 1882 (1 sp.)
- Doyerea Grosourdy 1864 (1 sp.)
- Eureiandra Hook. f. 1867 (8 spp.)
- Gurania (Schltdl.) Cogn. 1875 (37 spp.)
- Halosicyos Mart. Crov 1947 (1 sp.)
- Helmontia Cogn. 1875 (2–4 spp.)
- Ibervillea Greene 1895 (9–10 spp.)
- Kedrostis Medik. 1791 (28 spp.)
- Melotrianthus M. Crovetto 1954 (1–3 spp.)
- Psiguria Neck. ex Arn. 1841 (6–12 spp.)
- Seyrigia Keraudren 1960 [1961] (6 spp.)
- Trochomeriopsis Cogn. 1881 (1 sp.)
- Tumamoca Rose 1912 (2 spp.)
- Wilbrandia Silva Manso 1836 (5 spp.)
Tribu ti Benincaseae Ser. 1825
- Acanthosicyos Welw. ex Hook. f. 1867 (1 sp.)
- Benincasa Savi 1818 (2 spp., mairaman ti Praecitrullus Pangalo 1944)
- Borneosicyos (1–2 spp.)
- Cephalopentandra Chiov. 1929 (1 sp.)
- Citrullus Schrad. 1836 (4 spp.)
- Coccinia Wight & Arn. 1834 (30 spp.)
- Ctenolepis Hook. f. 1867 (3 spp.)
- Cucumis L. 1753 (65 spp.)
- Dactyliandra Hook. f. 1871 (2 spp.)
- Diplocyclos (Endl.) T. Post & Kuntze 1903 (4 spp.)
- Indomelothria (2 spp.)
- Khmeriosicyos (1 sp.)
- Lagenaria Ser. 1825 (6 spp.)
- Lemurosicyos Keraudren 1963 [1964] (1 sp.)
- Melothria L. 1753 (12 spp.)
- Muellerargia Cogn. 1881 (2 sp.)
- Papuasicyos (8 spp.)
- Peponium Engl. 1897 (20 spp.)
- Raphidiocystis Hook. f. 1867 (5 spp.)
- Ruthalicia C. Jeffrey 1962 (2 spp.)
- Scopellaria W.J. de Wilde & Duyfjes 2006 (2 spp.)
- Solena Lour. 1790 (3 spp.)
- Trochomeria Hook. f. 1867 (8 spp.)
- Zehneria Endl. 1833 (ca. 60 spp.)

Tribu ti Cucurbiteae Ser. 1825
- Abobra Naudin 1862 (1 sp.)
- Calycophysum H. Karst. & Triana 1854 [1855] (5 spp.)
- Cayaponia Silva Manso 1836 (50–59 spp., including Selysia Cogn. 1881)
- Cionosicys Griseb. 1864 [1860] (4–5 spp.)
- Cucurbita L. 1753 (15 spp.)
- Penelopeia Urb. 1921 (2 spp.)
- Peponopsis Naudin 1859 (1 sp.)
- Polyclathra Bertol. 1840 (6 spp.)
- Schizocarpum Schrad. 1830 (11 spp.)
- Sicana Naudin 1862 (4 spp.)
- Tecunumania Standl. & Steyerm. 1944 (1 sp.)

Alpabetiko a listaan dagiti henero

Abobra, Acanthosicyos, Actinostemma, Alsomitra, Ampelosicyos, Anisosperma, Apodanthera, Austrobryonia, Baijiania, Bambekea, Bayabusua, Benincasa, Borneosicyos, Bryonia, Calycophysum, Cayaponia, Cephalopentandra, Ceratosanthes, Cionosicys, Citrullus, Coccinia, Cogniauxia, Corallocarpus, Ctenolepis, Cucumis, Cucurbita, Cucurbitella, Cyclanthera, Cyclantheropsis, Dactyliandra, Dendrosicyos, Diplocyclos, Doyerea, Ecballium, Echinocystis, Echinopepon, Eureiandra, Fevillea, Frantzia, Gerrardanthus, Gomphogyne, Gurania, Gynostemma, Halosicyos, Hanburia, Helmontia, Hemsleya, Herpetospermum, Hodgsonia, Ibervillea, Indofevillea, Indomelothria, Kedrostis, Khmeriosicyos, Lagenaria, Lemurosicyos, Linnaeosicyos, Luffa, Marah, Melothria, Melotrianthus, Momordica, Muellerargia, Neoalsomitra, Nothoalsomitra, Papuasicyos, Penelopeia, Peponium, Peponopsis, Polyclathra, Psiguria, Pteropepon, Raphidiocystis, Ruthalicia, Schizocarpum, Schizopepon, Scopellaria, Seyrigia, Sicana, Sicydium, Sicyos, Siolmatra, Siraitia, Solena, Tecunumania, Telfairia, Thladiantha, Trichosanthes, Trochomeria, Trochomeriopsis, Tumamoca, Wilbrandia, Xerosicyos, Zanonia, Zehneria.

Sistematika

Mangisingasing ti moderno a molekular a pilohenetiko dagiti sumaganad a pannakikabagian:^[6]^[8]^[9]^[10]^[11]^[12]

Naisalaysayan a Kladograma a mangipakita ti pilohenio ti Cucurbitaceae

Cucurbitaceae

Gomphogyneae

Alsomitra

Bayabusua

	Hemsleya

	Gomphogyne

	Gynostemma

	Neoalsomitra

Triceratieae

Fevillea

Pteropepon

	Cyclantheropsis

	Sicydium

Zanonieae

Gerrardanthus

	Zanonia

	Siolmatra

Xerosicyos

Actinostemmateae	Actinostemma

Indofevilleeae	Indofevillea

Thladiantheae

	Baijiania

	Thladiantha

Siraitieae	Siraitia

Momordiceae	Momordica

Joliffieae

Cogniauxia

	Telfairia

	Ampelosicyos

Bryonieae

	Ecballium

	Bryonia

	Echinocystis

	Marah

	Echinopepon

	Frantzia

	Cyclanthera

	Hanburia

Sicyos

Linnaeosicyos

	Luffa

	Trichosanthes

Schizopeponeae

	Schizopepon

	Herpetospermum

Coniandreae

	Bambekea

	Eureiandra

Dendrosicyos

	Trochomeriopsis

	Seyrigia

Doyerea

	Helmontia

	Gurania

Melotrianthus

Kedrostis

	Ceratosanthes

	Halosicyos

Apodanthera

	Tumamoca

	Ibervillea

Cucurbiteae

Polyclathra

	Peponopsis

	Cucurbita

	Penelopeia

	Calycophysum

	Sicana

Tecunumania

Schizocarpum

Cionosicys

	Abobra

	Cayaponia

Benincaseae

Zehneria

Citrullus

	Peponium

	Lagenaria

	Solena

	Borneosicyos

Benincasa

Ctenolepis

	Dactyliandra

	Trochomeria

Ruthalicia

	Indomelothria

	Melothria

	Coccinia

	Diplocyclos

	Papuasicyos

	Scopellaria

	Muellerargia

	Cucumis

Dagiti nagibasaran

^ Angiosperm Phylogeny Group (2009). "An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants: APG III". Botanical Journal of the Linnean Society. 161 (2): 105–121. doi:10.1111/j.1095-8339.2009.00996.x.
^ Christenhusz, M. J. M. & Byng, J. W. (2016). "The number of known plants species in the world and its annual increase". Phytotaxa. 261 (3): 201–217. doi:10.11646/phytotaxa.261.3.1.
^ "Cucurbits". Purdue University. Naala idi Agosto 26, 2013.
^ "Angiosperm Phylogeny Website". mobot.org.
^ Revisions to Roland Brown's North American Paleocene Flora by Steven R. Manchester at Florida Museum of Natural History, University of Florida, Gainesville, Florida, USA. Published in Acta Musei Nationalis Pragae, Series B – Historia Naturalis, vol. 70, 2014, no. 3-4, pp. 153-210.
^ ^a ^b Schaefer H, Renner SS (2011). "Phylogenetic relationships in the order Cucurbitales and a new classification of the gourd family (Cucurbitaceae)". Taxon (journal). 60 (1): 122–138. JSTOR 41059827. Naiyarkibo manipud iti kasisigud idi 2018-11-06. Naala idi 2019-02-07.{{cite journal}}: Panagtaripato ti CS1: bot: di ammo ti kasasaad ti kasisigud nga URL (silpo)
^ Schaefer H, Kocyan A, Renner SS (2007). "Phylogenetics of Cucumis (Cucurbitaceae): Cucumber (C. sativus) belongs in an Asian/Australian clade far from melon (C. melo)". BMC Evolutionary Biology. 7: 58–69. doi:10.1186/1471-2148-7-58. PMC 3225884. PMID 17425784.
^ Zhang L-B, Simmons MP, Kocyan A, Renner SS (2006). "Phylogeny of the Cucurbitales based on DNA sequences of nine loci from three genomes: Implications for morphological and sexual system evolution" (PDF). Molecular Phylogenetics and Evolution. 39 (2): 305–322. doi:10.1016/j.ympev.2005.10.002. PMID 16293423. Naiyarkibo manipud iti kasisigud idi 27 February 2017.
^ Schaefer H, Heibl C, Renner SS (2009). "Gourds afloat: A dated phylogeny reveals an Asian origin of the gourd family (Cucurbitaceae) and numerous oversea dispersal events". Proceedings of the Royal Society B. 276 (1658): 843–851. doi:10.1098/rspb.2008.1447. PMC 2664369. PMID 19033142. Naiyarkibo manipud iti kasisigud idi 2 November 2018. Naala idi 7 February 2019.{{cite journal}}: Panagtaripato ti CS1: bot: di ammo ti kasasaad ti kasisigud nga URL (silpo)
^ de Boer HJ, Schaefer H, Thulin M, Renner SS (2012). "Evolution and loss of long-fringed petals: A case study using a dated phylogeny of the snake gourds, Trichosanthes (Cucurbitaceae)". BMC Evolutionary Biology. 12: 108. doi:10.1186/1471-2148-12-108. PMC 3502538. PMID 22759528.
^ Belgrano MJ (2012). Estudio sistemático y biogeográfico del género Apodanthera Arn. (Cucurbitaceae) [Systematic and biogeographic study of the genus Apodanthera Arn. (Cucurbitaceae)] (Ph.D.). Universidad Nacional de La Plata.
^ Renner SS, Schaefer H (2016). "Phylogeny and evolution of the Cucurbitaceae". Iti Grumet R, Katzir N, Garcia-Mas J (dagiti ed.). Genetics and Genomics of Cucurbitaceae. Plant Genetics and Genomics: Crops and Models. Vol. 20. New York, NY: Springer International Publishing. pp. 1–11. doi:10.1007/7397_2016_14. ISBN 978-3-319-49330-5.

Adu pay a mabasbasa

Bates D, Robinson R, Jeffrey C, dagiti ed. (1990). Biology and Utilization of the Cucurbitaceae. Cornell University Press. ISBN 978-0-8014-1670-5.
Jeffrey C. (2005). "A new system of Cucurbitaceae". Bot. Zhurn. 90: 332–335.

Dagiti akinruar a silpo

Dagiti midia a mainaig iti Cucurbitaceae iti Wikimedia Commons
Dagiti datos a mainaig iti Cucurbitaceae iti Wikispecies

[APGIII2009-1] Angiosperm Phylogeny Group (2009). "An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants: APG III". Botanical Journal of the Linnean Society. 161 (2): 105–121. doi:10.1111/j.1095-8339.2009.00996.x.

[Christenhusz-Byng2016-2] Christenhusz, M. J. M. & Byng, J. W. (2016). "The number of known plants species in the world and its annual increase". Phytotaxa. 261 (3): 201–217. doi:10.11646/phytotaxa.261.3.1.

[3] "Cucurbits". Purdue University. Naala idi Agosto 26, 2013.

[4] "Angiosperm Phylogeny Website". mobot.org.

[5] Revisions to Roland Brown's North American Paleocene Flora by Steven R. Manchester at Florida Museum of Natural History, University of Florida, Gainesville, Florida, USA. Published in Acta Musei Nationalis Pragae, Series B – Historia Naturalis, vol. 70, 2014, no. 3-4, pp. 153-210.

[Schaefer-6] Schaefer H, Renner SS (2011). "Phylogenetic relationships in the order Cucurbitales and a new classification of the gourd family (Cucurbitaceae)". Taxon (journal). 60 (1): 122–138. JSTOR 41059827. Naiyarkibo manipud iti kasisigud idi 2018-11-06. Naala idi 2019-02-07.{{cite journal}}: Panagtaripato ti CS1: bot: di ammo ti kasasaad ti kasisigud nga URL (silpo)

[7] Schaefer H, Kocyan A, Renner SS (2007). "Phylogenetics of Cucumis (Cucurbitaceae): Cucumber (C. sativus) belongs in an Asian/Australian clade far from melon (C. melo)". BMC Evolutionary Biology. 7: 58–69. doi:10.1186/1471-2148-7-58. PMC 3225884. PMID 17425784.

[8] Zhang L-B, Simmons MP, Kocyan A, Renner SS (2006). "Phylogeny of the Cucurbitales based on DNA sequences of nine loci from three genomes: Implications for morphological and sexual system evolution" (PDF). Molecular Phylogenetics and Evolution. 39 (2): 305–322. doi:10.1016/j.ympev.2005.10.002. PMID 16293423. Naiyarkibo manipud iti kasisigud idi 27 February 2017.

[9] Schaefer H, Heibl C, Renner SS (2009). "Gourds afloat: A dated phylogeny reveals an Asian origin of the gourd family (Cucurbitaceae) and numerous oversea dispersal events". Proceedings of the Royal Society B. 276 (1658): 843–851. doi:10.1098/rspb.2008.1447. PMC 2664369. PMID 19033142. Naiyarkibo manipud iti kasisigud idi 2 November 2018. Naala idi 7 February 2019.{{cite journal}}: Panagtaripato ti CS1: bot: di ammo ti kasasaad ti kasisigud nga URL (silpo)

[10] Boer HJ, Schaefer H, Thulin M, Renner SS (2012). "Evolution and loss of long-fringed petals: A case study using a dated phylogeny of the snake gourds, Trichosanthes (Cucurbitaceae)". BMC Evolutionary Biology. 12: 108. doi:10.1186/1471-2148-12-108. PMC 3502538. PMID 22759528.

[11] Belgrano MJ (2012). Estudio sistemático y biogeográfico del género Apodanthera Arn. (Cucurbitaceae) [Systematic and biogeographic study of the genus Apodanthera Arn. (Cucurbitaceae)] (Ph.D.). Universidad Nacional de La Plata.

[12] Renner SS, Schaefer H (2016). "Phylogeny and evolution of the Cucurbitaceae". Iti Grumet R, Katzir N, Garcia-Mas J (dagiti ed.). Genetics and Genomics of Cucurbitaceae. Plant Genetics and Genomics: Crops and Models. Vol. 20. New York, NY: Springer International Publishing. pp. 1–11. doi:10.1007/7397_2016_14. ISBN 978-3-319-49330-5.

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