Camponotus maculatus

Camponotus maculatus
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Hymenoptera
Family:	Formicidae
Subfamily:	Formicinae
Tribe:	Camponotini
Genus:	Camponotus
Species:	C. maculatus
Binomial name
Camponotus maculatus (Fabricius, 1782) Specimen labels
Subspecies
Camponotus maculatus foveolatus Stitz, 1925 Camponotus maculatus obfuscatus Viehmeyer, 1916 Camponotus maculatus subnudus Emery, 1889 Camponotus maculatus sylvaticomaculatus Dalla Torre, 1893 Camponotus maculatus ugandensis Santschi, 1923
Synonyms
Camponotus maculatus atramentarius Forel, 1904 Camponotus maculatus ballioni Forel, 1904 Camponotus maculatus boera Santschi, 1925 Camponotus maculatus cavallus Santschi, 1911 Camponotus maculatus cluisoides Forel, 1913 Camponotus maculatus conakryensis Emery, 1920 Camponotus maculatus erythraea Emery, 1920 Camponotus maculatus flavifemur Santschi, 1937 Camponotus maculatus flavominor Emery, 1925 Camponotus maculatus hannae Santschi, 1919 Camponotus maculatus hieroglyphicus Santschi, 1917 Camponotus maculatus intonsus Emery, 1905 Camponotus maculatus liocnemis Emery, 1905 Camponotus maculatus lividior Santschi, 1911 Camponotus maculatus lohieri Emery, 1915 Camponotus maculatus manzer Forel, 1910 Camponotus maculatus mathildae Forel, 1910 Camponotus maculatus melanocnemis Santschi, 1911 Camponotus maculatus miserabilis Santschi, 1914 Camponotus maculatus nubis Weber, 1943 Camponotus maculatus proletaria Baroni Urbani, 1971 Camponotus maculatus radamoide Forel, 1891 Camponotus maculatus sarmentus Emery, 1920 Camponotus maculatus schereri Forel, 1911 Camponotus maculatus schultzei Forel, 1912 Camponotus maculatus semispicatus Emery, 1920 Camponotus maculatus sudanicus Weber, 1943 Camponotus maculatus thomensis Santschi, 1920 Camponotus maculatus tuckeri Santschi, 1932 Camponotus maculatus zumpti Santschi, 1937 Camponotus sexpunctatus liengmei Forel, 1894 Formica cognata Smith, F., 1858 Formica lacteipennis Smith, F., 1858

Well known, common and widely distributed throughout Africa. A common ground nesting and foraging ant which often ascends trees to tend aphids, but does not build tents (Taylor 1977). It was not among the commoner species in the 76-farm survey of Taylor & Adedoyin (1978). Known to be prey of Oecophylla longinoda (Taylor et al., 2018). While this species has been suggested as being invasive in may parts of the world, these reports appear to be largely the result of misidentifications or are presented with little or no justification; additional details are required to confirm its occurrence outside its native range.

Identification

Keys including this Species

• Key to Camponotus maculatus species complex in the New World

Distribution

Latitudinal Distribution Pattern

Latitudinal Range: 49.78306° to -34.76667°.

North Temperate	North Subtropical	Tropical	South Subtropical	South Temperate

• Source: AntMaps

Distribution based on Regional Taxon Lists

Afrotropical Region: Angola, Benin, Cameroun, Central African Republic, Chad, Comoros, Eritrea, Ethiopia, Gambia, Guinea, Ivory Coast, Kenya, Lesotho, Liberia, Mali, Mozambique, Namibia, Sao Tome & Principe, Saudi Arabia, Senegal, Sierra Leone, South Africa, Sudan, Uganda, United Republic of Tanzania, Zimbabwe, Zimbabwe.
Indo-Australian Region: Micronesia (Federated States of), Palau.
Malagasy Region: Madagascar, Mauritius, Mayotte, Réunion.
Palaearctic Region: Algeria, Iran, Oman, Türkiye.

Distribution based on AntMaps

Distribution based on AntWeb specimens

Check data from AntWeb

Countries Occupied

Number of countries occupied by this species based on AntWiki Regional Taxon Lists. In general, fewer countries occupied indicates a narrower range, while more countries indicates a more widespread species.

Biology

Association with Other Organisms

 Explore: Show all Associate data or Search these data. See also a list of all data tables or learn how data is managed.

• This species is a mutualist for the aphid Cinara apini (a trophobiont) (Jones, 1927; Saddiqui et al., 2019).
• This species is a host for the eucharitid wasp Hydrorhoa striaticeps Kieffer complex (a parasitoid) (Quevillon, 2018) (multiple encounter modes; direct transmission; transmission outside nest).
• This species is a host for the eucharitid wasp Stilbula cyniformis (a parasite) (Universal Chalcidoidea Database) (primary host).
• This species is a host for the phorid fly Apocephalus coquilletti (a parasite) (phorid.net) (attacked).
• This species is a host for the strepsipteran Pseudacteon simplex (a parasite) (Quevillon, 2018) (encounter mode secondary; indirect transmission; transmission outside nest).

Castes

Worker

.	Owned by Museum of Comparative Zoology.

Images from AntWeb

Worker. Specimen code casent0055753. Photographer April Nobile, uploaded by California Academy of Sciences.	Owned by CAS, San Francisco, CA, USA.

Worker. Specimen code casent0063127. Photographer April Nobile, uploaded by California Academy of Sciences.	Owned by CAS, San Francisco, CA, USA.

Lectotype of Camponotus hova maculatoides. Worker. Specimen code casent0101095. Photographer April Nobile, uploaded by California Academy of Sciences.	Owned by NHMB, Basel, Switzerland.

Worker (major/soldier). Specimen code casent0101096. Photographer April Nobile, uploaded by California Academy of Sciences.	Owned by NHMB, Basel, Switzerland.

Paralectotype of Camponotus hova obscuratus. Worker (major/soldier). Specimen code casent0101109. Photographer April Nobile, uploaded by California Academy of Sciences.	Owned by NHMB, Basel, Switzerland.

Lectotype of Camponotus hova obscuratus. Worker. Specimen code casent0101110. Photographer April Nobile, uploaded by California Academy of Sciences.	Owned by NHMB, Basel, Switzerland.

Worker. Specimen code casent0101111. Photographer April Nobile, uploaded by California Academy of Sciences.	Owned by NHMB, Basel, Switzerland.

Worker. Specimen code casent0101182. Photographer April Nobile, uploaded by California Academy of Sciences.	Owned by NHMB, Basel, Switzerland.

Worker. Specimen code casent0101183. Photographer April Nobile, uploaded by California Academy of Sciences.	Owned by NHMB, Basel, Switzerland.

Paralectotype of Camponotus hova maculatoides. Worker (major/soldier). Specimen code casent0101338. Photographer April Nobile, uploaded by California Academy of Sciences.	Owned by MHNG, Geneva, Switzerland.

Paralectotype of Camponotus hova maculatoides. Worker. Specimen code casent0101339. Photographer April Nobile, uploaded by California Academy of Sciences.	Owned by MHNG, Geneva, Switzerland.

Worker. Specimen code casent0101340. Photographer Michele Esposito, uploaded by California Academy of Sciences.	Owned by MHNG, Geneva, Switzerland.

Worker (major/soldier). Specimen code casent0101341. Photographer Michele Esposito, uploaded by California Academy of Sciences.	Owned by MHNG, Geneva, Switzerland.

Worker (major/soldier). Specimen code casent0101352. Photographer April Nobile, uploaded by California Academy of Sciences.	Owned by MHNG, Geneva, Switzerland.

Worker. Specimen code casent0101353. Photographer April Nobile, uploaded by California Academy of Sciences.	Owned by MHNG, Geneva, Switzerland.

Worker (major/soldier). Specimen code casent0101354. Photographer April Nobile, uploaded by California Academy of Sciences.	Owned by MHNG, Geneva, Switzerland.

Worker. Specimen code casent0101356. Photographer April Nobile, uploaded by California Academy of Sciences.	Owned by MHNG, Geneva, Switzerland.

Paralectotype of Camponotus hova maculatoides. Worker (major/soldier). Specimen code casent0101747. Photographer April Nobile, uploaded by California Academy of Sciences.	Owned by MHNG, Geneva, Switzerland.

Worker (major/soldier). Specimen code casent0101929. Photographer April Nobile, uploaded by California Academy of Sciences.	Owned by MHNG, Geneva, Switzerland.

Worker (major/soldier). Specimen code casent0103332. Photographer April Nobile, uploaded by California Academy of Sciences.	Owned by CAS, San Francisco, CA, USA.

Worker. Specimen code casent0055750. Photographer April Nobile, uploaded by California Academy of Sciences.	Owned by CAS, San Francisco, CA, USA.

Syntype of Camponotus grandidieri comorensis. Worker. Specimen code casent0104631. Photographer April Nobile, uploaded by California Academy of Sciences.	Owned by ZMHB, Berlin, Germany.

Paratype of Camponotus rubripes hagensii. Worker. Specimen code casent0104633. Photographer April Nobile, uploaded by California Academy of Sciences.	Owned by ZMHB, Berlin, Germany.

Type of unavailable quadrinomial: Camponotus maculatus hova laticollus. Worker (major/soldier). Specimen code casent0104638. Photographer April Nobile, uploaded by California Academy of Sciences.	Owned by ZMHB, Berlin, Germany.

Worker. Specimen code casent0135006. Photographer April Nobile, uploaded by California Academy of Sciences.	Owned by CAS, San Francisco, CA, USA.

Worker (major/soldier). Specimen code casent0146442. Photographer Erin Prado, uploaded by California Academy of Sciences.	Owned by CAS, San Francisco, CA, USA.

Worker. Specimen code casent0179509. Photographer Erin Prado, uploaded by California Academy of Sciences.	Owned by CAS, San Francisco, CA, USA.

Queen

Images from AntWeb

Queen (alate/dealate). Specimen code casent0147206. Photographer Erin Prado, uploaded by California Academy of Sciences.	Owned by CAS, San Francisco, CA, USA.

Queen (alate/dealate). Specimen code casent0125148. Photographer April Nobile, uploaded by California Academy of Sciences.	Owned by CAS, San Francisco, CA, USA.

Queen (alate/dealate). Specimen code casent0101184. Photographer April Nobile, uploaded by California Academy of Sciences.	Owned by NHMB, Basel, Switzerland.

Queen (alate/dealate). Specimen code casent0055751. Photographer April Nobile, uploaded by California Academy of Sciences.	Owned by CAS, San Francisco, CA, USA.

Queen (alate/dealate). Specimen code casent0101879. Photographer April Nobile, uploaded by California Academy of Sciences.	Owned by MHNG, Geneva, Switzerland.

Male

Images from AntWeb

Male (alate). Specimen code casent0101185. Photographer April Nobile, uploaded by California Academy of Sciences.	Owned by NHMB, Basel, Switzerland.

Male (alate). Specimen code casent0101569. Photographer April Nobile, uploaded by California Academy of Sciences.	Owned by MHNG, Geneva, Switzerland.

Male (alate). Specimen code casent0146792. Photographer Erin Prado, uploaded by California Academy of Sciences.	Owned by CAS, San Francisco, CA, USA.

Male (alate). Specimen code casent0101355. Photographer April Nobile, uploaded by California Academy of Sciences.	Owned by MHNG, Geneva, Switzerland.

Nomenclature

The following information is derived from Barry Bolton's Online Catalogue of the Ants of the World.

• maculatus. Formica maculata Fabricius, 1782: 491 (w.) “Africa”.
  • Type-material: holotype major worker.
  • Type-locality: Africa: (“Habitat in Africa aequinoctiali. Mus. Dom. Banks.”) (no further data).
  • [Note: Donisthorpe, 1915a: 221, says the type-locality is “probably Sierra Leone, or nearby on the West Coast”.]
  • Type-depository: BMNH.
  • Smith, F. 1858b: 29 (s.w.); Mayr, 1862: 654 (q.m.); Forel, 1891b: 29 (s.w.q.m.).
  • Combination in Camponotus: Mayr, 1862: 654;
  • combination in C. (Myrmoturba): Forel, 1912i: 91;
  • combination in C. (Tanaemyrmex): Emery, 1925b: 86.
  • Subspecies of sylvaticus: Forel, 1879a: 64; Emery & Forel, 1879: 448; Mayr, 1880: 21; Emery, 1881b: 526; André, 1882a: 145; André, 1884b: 536; Emery, 1884a: 375; Emery, 1887a: 212 (footnote).
  • Subspecies of rubripes: Forel, 1886f: 143, 149; Emery, 1887a: 212 (footnote).
  • Status as species: Fabricius, 1787: 308; Gmelin, 1790: 2800; Olivier, 1792: 495; Fabricius, 1793: 356; Latreille, 1802c: 283; Fabricius, 1804: 403; Lepeletier de Saint-Fargeau, 1835: 214; Haliday, 1836: 329; Smith, F. 1858b: 28 (redescription); Gerstäcker, 1859: 262; Mayr, 1862: 654 (redescription); Gerstäcker, 1862: 509; Mayr, 1863: 400; Roger, 1863b: 2; Mayr, 1865: 27; Gerstäcker, 1871: 355; Mayr, 1877: 20 (in list); Emery, 1877b: 364; Emery, 1878b: 44; Forel, 1891b: 23, 29 (redescription); Emery, 1891b: 17; Forel, 1892e: 349; Emery, 1892a: 119; Emery, 1892d: 566; Dalla Torre, 1893: 241; Emery, 1893i: 671; Mayr, 1893: 195; Mayr, 1895: 150; Emery, 1895a: 180; Emery, 1895h: 46; Emery, 1896d: 370 (in list); Emery, 1896i: 158; Mayr, 1896: 249; Mayr, 1897: 432; Emery, 1897e: 604; Emery, 1898a: 225; Forel, 1899c: 134; Emery, 1899a: 501; Mayr, 1901b: 28; Mayr, 1904b: 11; Emery, 1905d: 37; Ruzsky, 1905b: 191; Mayr, 1907a: 389; Mayr, 1907b: 19; Forel, 1907e: 16; Forel, 1907g: 88; Forel, 1908a: 16; Emery, 1908a: 188 (infraspecific taxa key); Santschi, 1908: 518; Bondroit, 1910: 487; Forel, 1910c: 266; Forel, 1910f: 26; Karavaiev, 1910b: 9; Stitz, 1910: 148; Wheeler, W.M. 1910d: 301; Stitz, 1911b: 392; Forel, 1911d: 371; Viehmeyer, 1912: 8; Forel, 1913a: 145; Forel, 1913b: 340; Santschi, 1913h: 42; Santschi, 1914b: 129; Santschi, 1914c: 288; Santschi, 1914d: 379; Forel, 1914d: 249; Wheeler, W.M. & Mann, 1914: 47; Donisthorpe, 1915a: 221 (redescription of holotype); Wheeler, W.M. 1915g: 814; Forel, 1915d: 70 (in key); Emery, 1915f: 79; Emery, 1916b: 224; Stitz, 1916: 403; Mann, 1916: 474; Stitz, 1917: 351; Luederwaldt, 1918: 49; Mann, 1920: 435; Emery, 1920c: 5, 12; Arnold, 1922: 621; Wheeler, W.M. 1922a: 234, 949, 1040; Soudek, 1922: 91; Viehmeyer, 1923: 93; Stitz, 1923: 164; Wheeler, W.M. 1924b: 253; Wheeler, W.M. 1925b: 3; Emery, 1925b: 86; Santschi, 1925h: 166; Crawley, 1926: 393 (redescription); Menozzi, 1926a: 41; Karavaiev, 1926e: 189; Arnold, 1926: 292; Karavaiev, 1927a: 294; Santschi, 1932f: 519; Finzi, 1936: 189; Santschi, 1936e: 81; Finzi, 1939a: 159; Finzi, 1940: 163; Weber, 1943c: 385; Donisthorpe, 1946e: 33; Chapman & Capco, 1951: 248; Bernard, 1953b: 260; Prins, 1963: 106; Baroni Urbani, 1972: 125; Hamann & Klemm, 1976: 676; Collingwood, 1985: 282; Marsh, 1986: 342; Bolton, 1995b: 109; Collingwood & Agosti, 1996: 374; Blard, et al. 2003: 129; Paknia, et al. 2008: 153; Mackay & Delsinne, 2009: 496 (in key); Legakis, 2011: 31; Borowiec, L. & Salata, 2012: 480; Hita Garcia, et al. 2013: 205; Borowiec, L. 2014: 36 (see note in bibliography); Madl, 2019: 14.
  • Senior synonym of atramentarius: Baroni Urbani, 1972: 126; Bolton, 1995b: 110.
  • Senior synonym of ballioni: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
  • Senior synonym of boera: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
  • Senior synonym of cavallus: Baroni Urbani, 1972: 126; Bolton, 1995b: 110.
  • Senior synonym of cluisoides: Baroni Urbani, 1972: 124; Bolton, 1995b: 110.
  • Senior synonym of cognata: Baroni Urbani, 1972: 127; Bolton, 1995b: 110.
  • Senior synonym of conakryensis: Baroni Urbani, 1972: 124; Bolton, 1995b: 110.
  • Senior synonym of erythraea: Baroni Urbani, 1972: 124; Bolton, 1995b: 110.
  • Senior synonym of flavifemur: Baroni Urbani, 1972: 124; Bolton, 1995b: 110.
  • Senior synonym of flavominor: Baroni Urbani, 1972: 124; Bolton, 1995b: 110.
  • Senior synonym of hannae: Baroni Urbani, 1972: 126; Bolton, 1995b: 110.
  • Senior synonym of hieroglyphicus: Baroni Urbani, 1972: 126; Bolton, 1995b: 110.
  • Senior synonym of intonsus: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
  • Senior synonym of lacteipennis: Baroni Urbani, 1972: 127; Bolton, 1995b: 110.
  • Senior synonym of liengmei: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
  • Senior synonym of liocnemis: Baroni Urbani, 1972: 126; Bolton, 1995b: 110.
  • Senior synonym of lividior: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
  • Senior synonym of lohieri: Baroni Urbani, 1972: 124; Bolton, 1995b: 110.
  • Senior synonym of manzer: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
  • Senior synonym of mathildae: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
  • Senior synonym of melanocnemis: Baroni Urbani, 1972: 124; Bolton, 1995b: 110.
  • Senior synonym of miserabilis: Baroni Urbani, 1972: 126; Bolton, 1995b: 110.
  • Senior synonym of nubis: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
  • Senior synonym of proletaria: Baroni Urbani, 1972: 126; Bolton, 1995b: 110.
  • Senior synonym of radamoides: Baroni Urbani, 1972: 126; Bolton, 1995b: 110.
  • Senior synonym of sarmentus: Baroni Urbani, 1972: 126; Bolton, 1995b: 110.
  • Senior synonym of schereri: Baroni Urbani, 1972: 126; Bolton, 1995b: 110.
  • Senior synonym of schultzei: Baroni Urbani, 1972: 124; Bolton, 1995b: 110.
  • Senior synonym of semispicatus: Baroni Urbani, 1972: 124; Bolton, 1995b: 110.
  • Senior synonym of sudanicus: Baroni Urbani, 1972: 126; Bolton, 1995b: 110.
  • Senior synonym of thomensis: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
  • Senior synonym of tuckeri: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
  • Senior synonym of zumpti: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
  • Material of the unavailable names calceatus, cataractae, citinus, cluis, contaminatus, diffusus, hansingi, importunoides, madecassa, pessimus referred here by Baroni Urbani, 1972: 124-127; Bolton, 1995b: 110.
  • Distribution:
  • Afrotropical: Angola, Benin, Botswana, Cameroon, Congo, Democratic Republic of Congo, Equatorial Guinea (+ Bioko), Eritrea, Ethiopia, Gambia, Ghana, Guinea, Guinea Bissau, Ivory Coast, Kenya, Liberia, Mozambique, Namibia, Nigeria, São Tomé & Principe, Senegal, Sierra Leone, Somalia, South Africa, South Sudan, Sudan, Tanzania (+ Zanzibar), Togo, Uganda, Zambia, Zimbabwe.
  • Malagasy: Comoros, Madagascar.
  • Palaearctic: Egypt, Iran, Oman, Tunisia, Turkey, Turkmenistan, Yemen.
  • Current subspecies: nominal plus foveolatus (unresolved junior homonym), obfuscatus, subnudus, sylvaticomaculatus, ugandensis.
• atramentarius. Camponotus maculatus var. atramentarius Forel, 1904b: 379 (w.) ETHIOPIA.
  • Type-material: holotype (?) worker.
  • [Note: no indication of number of specimens is given.]
  • Type-locality: Ethiopia (“Abyssinia”): (no further data) (Dmitriev).
  • Type-depository: ZISP (perhaps also MHNG).
  • Forel, 1910f: 26 (q.).
  • Combination in C. (Myrmoturba): Forel, 1913b: 340;
  • combination in C. (Tanaemyrmex): Emery, 1925b: 88.
  • Subspecies of maculatus: Forel, 1905b: 180; Forel, 1907e: 14; Forel, 1907g: 88; Forel, 1909b: 54, 67; Forel, 1910f: 26; Forel, 1911d: 371; Forel, 1911f: 280; Forel, 1913b: 340; Santschi, 1914b: 131; Emery, 1920c: 5; Arnold, 1922: 625; Wheeler, W.M. 1922a: 952; Stitz, 1923: 164; Emery, 1925b: 88; Santschi, 1930b: 75; Santschi, 1937d: 240; Bernard, 1953b: 261.
  • Junior synonym of maculatus: Baroni Urbani, 1972: 126; Bolton, 1995b: 86.
• ballioni. Camponotus maculatus r. ballioni Forel, 1904d: 176 (s.w.q.) SOUTH AFRICA.
  • Type-material: syntype major and minor workers, syntype queen(s) (numbers not stated).
  • Type-localities: South Africa: Cape (ex coll. Ballion, ex coll. Puls) (invalid restriction of type-locality by Wheeler, W.M. 1922a: 954; no lectotype designated), South Africa: Port Elisabeth (H. Brauns).
  • Type-depository: MRHN (probably also MHNG).
  • Combination in C. (Tanaemyrmex): Emery, 1925b: 87.
  • As unavailable (infrasubspecific) name: Wheeler, W.M. 1922a: 954.
  • Status as species: Santschi, 1925h: 166.
  • Subspecies of maculatus: Emery, 1920c: 5; Emery, 1925b: 87; Santschi, 1926b: 249.
  • Junior synonym of maculatus: Baroni Urbani, 1972: 125; Bolton, 1995b: 87.
• boera. Camponotus (Myrmoturba) ballioni var. boera Santschi, 1925h: 166.
  • [First available use of Camponotus maculatus subsp. cognatus var. boera Forel, 1910f: 27 (w.q.) SOUTH AFRICA; unavailable (infrasubspecific) name.]
  • Type-material: syntype workers (number not stated), 1 syntype queen.
  • Type-localities: South Africa: Cape Prov., “Klein Namaland”, Steinkopf (L. Schultze) (invalid restriction of type-locality by Wheeler, W.M. 1922a: 954; no lectotype designated), South Africa: “Klein Namaland”, Kamaggas (L. Schultze).
  • Type-depository: MHNG.
  • As unavailable (infrasubspecific) name: Emery, 1920c: 5; Arnold, 1922: 632; Wheeler, W.M. 1922a: 954; Emery, 1925b: 87.
  • Subspecies of ballioni: Santschi, 1930b: 57.
  • Junior synonym of maculatus: Baroni Urbani, 1972: 125; Bolton, 1995b: 89.
• cavallus. Camponotus maculatus st. cavallus Santschi, 1911g: 211 (s.w.q.) ANGOLA.
  • Type-material: syntype major and minor workers, syntype queens (numbers not stated).
  • Type-locality: Angola: Benguela, Cucala (J. Cruchet).
  • Type-depository: NHMB.
  • Combination in C. (Tanaemyrmex): Emery, 1925b: 88.
  • Subspecies of maculatus: Stitz, 1916: 403; Emery, 1920c: 5; Wheeler, W.M. 1922a: 953; Emery, 1925b: 88; Santschi, 1930b: 57; Santschi, 1937d: 241.
  • Junior synonym of maculatus: Baroni Urbani, 1972: 126; Bolton, 1995b: 91.
• cluisoides. Camponotus (Myrmoturba) maculatus var. cluisoides Forel, 1913h: 354 (s.w.) KENYA.
  • Type-material: syntype major and minor workers (number not stated).
  • Type-locality: Kenya (“Afrique orientale anglaise”): Tsavo, Campo Tembo (Bayer).
  • Type-depository: MRAC (perhaps also MHNG).
  • Combination in C. (Tanaemyrmex): Emery, 1925b: 87.
  • As unavailable (infrasubspecific) name: Emery, 1920c: 5; Wheeler, W.M. 1922a: 952.
  • Subspecies of maculatus: Emery, 1925b: 87; Menozzi, 1927f: 361.
  • Junior synonym of maculatus: Baroni Urbani, 1972: 124; Bolton, 1995b: 93.
• cognata. Formica cognata Smith, F. 1858b: 35 (s.w.) SOUTH AFRICA.
  • Type-material: syntype major and minor workers (numbers not stated).
  • Type-locality: South Africa: Port Natal (= Durban) (no collector’s name).
  • Type-depository: BMNH.
  • Mayr, 1862: 655 (q.m.).
  • Combination in Camponotus: Mayr, 1862: 655;
  • combination in C. (Tanaemyrmex): Emery, 1925b: 88.
  • As unavailable (infrasubspecific) name: Emery, 1894i: 46; Emery, 1895h: 46.
  • Status as species: Mayr, 1877: 20 (in list); Dalla Torre, 1893: 225.
  • Subspecies of sylvaticus: Forel, 1879a: 65; Emery & Forel, 1879: 448; Mayr, 1880: 22; Emery, 1881b: 526; André, 1882a: 146; Emery, 1882: 448; Emery, 1884a: 375; André, 1884b: 536; Forel, 1885b: 176; Forel, 1886d: 239; Forel, 1886e: clxvii; Medina, 1893: 104.
  • Subspecies of rubripes: Forel, 1886f: 143, 151; Emery, 1887a: 212 (footnote); Mayr, 1889: 278; Forel, 1889: 255; Forel, 1890a: lxv; Saunders, E. 1890: 202.
  • Subspecies of maculatus: Mayr, 1862: 655; Emery, 1891b: 19; Emery, 1892d: 566; Emery, in Dalla Torre, 1893: 225, 241 (footnote); Mayr, 1893: 195; André, 1895a: 1; Mayr, 1896: 249; Forel, 1904b: 378; Forel, 1904c: 12; Ruzsky, 1905b: 205; Stitz, 1910: 148; Forel, 1910f: 25; Emery, 1920c: 5; Wheeler, W.M. 1922a: 953; Arnold, 1922: 631; Emery, 1925b: 88; Menozzi, 1942: 180; Eidmann, 1944: 469.
  • Junior synonym of maculatus: Roger, 1863b: 2; Mayr, 1863: 400; Mayr, 1865: 27; Gerstäcker, 1871: 355; Baroni Urbani, 1972: 127; Bolton, 1995b: 93.
• conakryensis. Camponotus (Myrmoturba) maculatus var. conakryensis Emery, 1920c: 13 (s.w.) GUINEA.
  • Type-material: syntype major and minor workers (numbers not stated).
  • Type-locality: Guinea (“Guinée française”): Conakry (F. Silvestri).
  • Type-depository: MSNG.
  • Subspecies of maculatus: Emery, 1925b: 87; Santschi, 1930a: 73.
  • Junior synonym of maculatus: Baroni Urbani, 1972: 124; Bolton, 1995b: 93.
• erythraea. Camponotus (Myrmoturba) maculatus var. erythraea Emery, 1920c: 14 (s.) ERITREA.
  • Type-material: holotype major worker.
  • Type-locality: Eritrea: Asmara (F. Silvestri).
  • Type-depository: MSNG.
  • Subspecies of maculatus: Emery, 1925b: 87; Santschi, 1935b: 275.
  • Junior synonym of maculatus: Baroni Urbani, 1972: 124; Bolton, 1995b: 98.
• flavifemur. Camponotus (Myrmoturba) maculatus var. flavifemur Santschi, 1937g: 84 (s.w.) TANZANIA.
  • Type-material: 1 syntype major worker, 1 syntype minor worker.
  • Type-locality: Tanzania: Lake Victoria, Mwanga, vii.-viii.1931 (M. Grabham).
  • Type-depository: BMNH.
  • Junior synonym of maculatus: Baroni Urbani, 1972: 124; Bolton, 1995b: 99.
• flavominor. Camponotus (Tanaemyrmex) maculatus var. flavominor Emery, 1925b: 87.
  • [First available use of Camponotus (Myrmoturba) maculatus st. melanocnemis var. flavominor Santschi, 1920i: 4 (s.w.) BENIN (footnote); unavailable (infrasubspecific) name.]
  • Type-material: syntype major and minor workers (numbers not stated).
  • Type-locality: Benin (“Upper Dahomey”): Kouandé (Desanti).
  • Type-depository: NHMB.
  • As unavailable (infrasubspecific) name: Wheeler, W.M. 1922a: 956.
  • Junior synonym of maculatus: Baroni Urbani, 1972: 124; Bolton, 1995b: 100.
• hannae. Camponotus (Myrmoturba) maculatus st. hannae Santschi, 1919a: 349, fig. 4 (s.w.) ZIMBABWE.
  • Type-material: syntype major and minor workers (numbers not stated).
  • Type-locality: Zimbabawe (“Rhodesia”): Victoria Falls (G. Arnold).
  • Type-depository: NHMB.
  • Combination in C. (Tanaemyrmex): Emery, 1925b: 88.
  • Subspecies of maculatus: Emery, 1920c: 5; Wheeler, W.M. 1922a: 955; Emery, 1925b: 88; Arnold, 1926: 294.
  • Junior synonym of maculatus: Baroni Urbani, 1972: 126; Bolton, 1995b: 102.
• hieroglyphicus. Camponotus maculatus st. hieroglyphicus Santschi, 1917b: 290 (s.w.q.) ANGOLA.
  • Type-material: syntype major and minor workers, syntype queen(s) (numbers not stated).
  • Type-locality: Angola: Benguela, Caconda (J. Cruchet).
  • Type-depository: NHMB.
  • Combination in C. (Tanaemyrmex): Emery, 1925b: 88.
  • Subspecies of maculatus: Emery, 1920c: 5; Wheeler, W.M. 1922a: 955; Emery, 1925b: 88.
  • Junior synonym of maculatus: Baroni Urbani, 1972: 126; Bolton, 1995b: 103.
• intonsus. Camponotus maculatus subsp. intonsus Emery, 1905d: 29 (footnote) (s.w.) SOUTH AFRICA.
  • Type-material: syntype major and minor workers (numbers not stated).
  • Type-locality: South Africa: Cape Prov. (no collector’s name) (invalid restriction of type-locality by Wheeler, W.M. 1922a: 955; no lectotype designated) South Africa: Transvaal (no collector’s name).
  • Type-depository: MSNG.
  • As unavailable (infrasubspecific) name: Emery, 1920c: 5; Emery, 1925b: 87; Santschi, 1926b: 249.
  • Subspecies of maculatus: Arnold, 1922: 632; Wheeler, W.M. 1922a: 955.
  • Junior synonym of maculatus: Baroni Urbani, 1972: 125; Bolton, 1995b: 105.
• lacteipennis. Formica lacteipennis Smith, F. 1858b: 34 (s.w.q.m.) SOUTH AFRICA.
  • Type-material: syntype major and minor workers, syntype queen(s), syntype male(s) (numbers not stated).
  • Type-locality: South Africa: Port Natal (= Durban) (R.W. Plant).
  • Type-depository: BMNH.
  • Combination in Camponotus: Roger, 1863b: 2;
  • combination in C. (Myrmoturba): Forel, 1914a: 267;
  • combination in C. (Tanaemyrmex): Emery, 1925b: 90.
  • Status as species: Mayr, 1863: 416; Roger, 1863b: 2; Dalla Torre, 1893: 237.
  • Subspecies of sylvaticus: Mayr, 1886c: 355.
  • Subspecies of maculatus: Emery, 1895h: 46; Emery, 1896d: 370 (in list); Emery, 1920c: 5; Arnold, 1922: 626; Wheeler, W.M. 1922a: 955; Emery, 1925b: 89.
  • Junior synonym of maculatus: Baroni Urbani, 1972: 127; Bolton, 1995b: 107.
• liengmei. Camponotus sexpunctatus var. liengmei Forel, 1894b: 67 (s.w.) MOZAMBIQUE.
  • Type-material: syntype major and minor workers (numbers not stated).
  • Type-locality: Mozambique: Delagoa Bay (Liengme).
  • Type-depository: MHNG.
  • Forel, 1907g: 88 (m.).
  • Combination in C. (Myrmoturba): Forel, 1914a: 267;
  • combination in C. (Dinomyrmex): Santschi, 1921f: 312;
  • combination in C. (Tanaemyrmex): Emery, 1925b: 87.
  • As unavailable (infrasubspecific) name: Emery, 1895h: 46; Emery, 1896d: 371 (in list.
  • Subspecies of maculatus: Forel, 1901h: 69; Forel, 1907g: 88; Forel, 1910e: 452; Forel, 1911e: 288; Forel, 1914d: 249; Emery, 1920c: 5; Arnold, 1922: 636; Wheeler, W.M. 1922a: 955; Emery, 1925b: 87; Prins, 1963: 107.
  • Status as species: Santschi, 1923e: 292.
  • Junior synonym of maculatus Baroni Urbani, 1972: 125; Bolton, 1995b: 108.
• liocnemis. Camponotus maculatus subsp. liocnemis Emery, 1905d: 30 (footnote) (s.w.) SOUTH AFRICA.
  • Type-material: syntype major and minor workers (numbers not stated).
  • Type-localities: South Africa: Transvaal (no collector’s name) (invalid restriction of type-locality by Wheeler, W.M. 1922a: 958; no lectotype designated), South Africa: Cape Prov. (no collector’s name).
  • [Note: Wheeler, W.M. 1922a: 958, provides the syntype data Transvaal, Makapan (E. Simon).]
  • Type-depository: MSNG.
  • [Misspelled as lioculmis by Forel, 1905b: 180 (in text); misspelled as liochnemis by Santschi, 1914e: 38.]
  • Santschi, 1914e: 38 (q.m.).
  • Combination in C. (Myrmoturba): Santschi, 1914d: 379.
  • As unavailable (infrasubspecific) name: Forel, 1907e: 16; Forel, 1909b: 54; Forel, 1910f: 26; Forel, 1911d: 371; Forel, 1913b: 340; Santschi, 1917b: 293; Emery, 1920c: 5; Arnold, 1922: 625; Wheeler, W.M. 1922a: 958; Emery, 1925b: 88; Santschi, 1925h: 166; Santschi, 1930a: 73; Santschi, 1930b: 76.
  • Subspecies of maculatus: Santschi, 1914d: 379; Santschi, 1914e: 38.
  • Junior synonym of maculatus: Baroni Urbani, 1972: 126; Bolton, 1995b: 108.
• lividior. Camponotus maculatus var. lividior Santschi, 1911e: 128 (s.w.q.m.) COMOROS.
  • Type-material: syntype major and minor workers, syntype queen(s), syntype male(s) (numbers not stated).
  • Type-locality: Comoros: Grande Comoro I., 1899 (H. Pobequin).
  • Type-depositories: MNHN, NHMB.
  • Subspecies of maculatus: Emery, 1920c: 5; Emery, 1925b: 88.
  • Junior synonym of maculatus: Baroni Urbani, 1972: 125; Bolton, 1995b: 108.
• lohieri. Camponotus (Myrmoturba) maculatus var. lohieri Emery, 1915g: 22.
  • [First available use of Camponotus (Myrmoturba) maculatus st. melanocnemis var. lohieri Santschi, 1913c: 313 (w.) IVORY COAST; unavailable (infrasubspecific) name.]
  • Type-material: holotype (?) worker.
  • [Note: no indication of number of specimens is given.]
  • Type-locality: Ivory Coast: Jacqueville (J.H. Lohier).
  • Type-depository: NHMB.
  • As unavailable (infrasubspecific) name: Santschi, 1914d: 379; Santschi, 1915c: 279; Emery, 1920c: 5; Wheeler, W.M. 1922a: 957.
  • Subspecies of maculatus: Emery, 1925b: 87; Menozzi, 1942: 180; Eidmann, 1944: 463.
  • Junior synonym of maculatus: Baroni Urbani, 1972: 124; Bolton, 1995b: 109.
• manzer. Camponotus maculatus subsp. manzer Forel, 1910e: 452 (s.) ZIMBABWE.
  • Type-material: holotype major worker.
  • Type-locality: Zimbabwe: Zambezi, Victoria Falls (R.C. Wroughton).
  • Type-depository: MHNG.
  • Forel, 1911f: 280 (w.); Santschi, 1916b: 509 (m.).
  • Combination in C. (Myrmoturba): Santschi, 1916b: 509;
  • combination in C. (Tanaemyrmex): Emery, 1925b: 88.
  • Subspecies of maculatus: Forel, 1911f: 280; Santschi, 1916b: 509; Santschi, 1917b: 291; Emery, 1920c: 5; Arnold, 1922: 624; Wheeler, W.M. 1922a: 956; Emery, 1925b: 88; Santschi, 1930b: 76.
  • Junior synonym of maculatus: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
• mathildae. Camponotus maculatus subsp. mathildae Forel, 1910c: 266 (s.w.q.) SOUTH AFRICA.
  • Type-material: syntype major and minor workers, syntype queen(s) (numbers not stated).
  • Type-locality: South Africa: Transvaal, Zoutpansberg, 800 m. (no collector’s name) (received from Rolle).
  • Type-depository: MHNG.
  • Combination in C. (Myrmoturba): Santschi, 1914b: 130;
  • combination in C. (Tanaemyrmex): Emery, 1925b: 88.
  • As unavailable (infrasubspecific) name: Arnold, 1922: 649.
  • Subspecies of maculatus: Santschi, 1914b: 130; Emery, 1920c: 5; Wheeler, W.M. 1922a: 956; Emery, 1925b: 88.
  • Junior synonym of maculatus: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
• melanocnemis. Camponotus maculatus var. melanocnemis Santschi, 1911c: 368 (s.) DEMOCRATIC REPUBLIC OF CONGO.
  • Type-material: syntype major workers (number not stated).
  • Type-localities: Congo (“Congo français”): (no further data), 1894 (H. Pobeguin) (invalid restriction of type-locality by Wheeler, W.M. 1922a: 956; no lectotype designated), Congo (“Congo français”): Grand Lahou, 1894 (H. Pobeguin).
  • Type-depository: MNHN.
  • Santschi, 1915c: 278 (w.).
  • As unavailable (infrasubspecific) name: Emery, 1920c: 5.
  • Subspecies of maculatus: Forel, 1913h: 355; Santschi, 1913c: 313; Forel, 1914d: 250; Santschi, 1914d: 379; Santschi, 1915c: 278; Santschi, 1920i: 4 (footnote); Wheeler, W.M. 1922a: 234, 956; Emery, 1925b: 87; Weber, 1943c: 333; Bernard, 1953b: 260.
  • Junior synonym of maculatus: Baroni Urbani, 1972: 124; Bolton, 1995b: 111.
• miserabilis. Camponotus (Myrmoturba) maculatus st. miserabilis Santschi, 1914d: 379 (s.w.) GUINEA.
  • Type-material: 4 syntype major and minor workers.
  • Type-locality: Guinea (“Guinée française”): Kindia (F. Silvestri).
  • Type-depository: NHMB.
  • [Junior primary homonym of *Camponotus miserabilis Förster, 1891: 430.]
  • Subspecies of maculatus: Santschi, 1915c: 281; Emery, 1920c: 5; Wheeler, W.M. 1922a: 219, 957; Emery, 1925b: 88.
  • Replacement name: Camponotus maculatus subsp. proletaria Baroni Urbani, 1971b: 362.
  • Junior synonym of maculatus: Baroni Urbani, 1972: 126.
• nubis. Camponotus (Myrmoturba) maculatus subsp. nubis Weber, 1943c: 385, pl. 16, fig. 30 (s.w.) SOUTH SUDAN.
  • Type-material: syntype major and minor workers (numbers not stated, “one colony”).
  • Type-locality: South Sudan (“Anglo-Egyptian Sudan”): Imatong Mts, 8700 ft, 28.vii.1939, no. 1351 (N.A. Weber).
  • Type-depository: MCZC.
  • Junior synonym of maculatus: Baroni Urbani, 1972: 125; Bolton, 1995b: 114.
• proletaria. Camponotus maculatus subsp. proletaria Baroni Urbani, 1971b: 362.
  • Replacement name for Camponotus (Myrmoturba) maculatus st. miserabilis Santschi, 1914d: 379. [Junior primary homonym of *Camponotus miserabilis Förster, 1891: 430.]
  • Junior synonym of maculatus: Baroni Urbani, 1972: 126; Bolton, 1995b: 118.
• radamoides Camponotus maculatus var. radamoides Forel, 1891b: 213 (in text) (s.w.) MADAGASCAR.
  • Type-material: syntype major and minor workers (numbers not stated).
  • Type-locality: Madagascar: Forêt d’Andrangoloaka (F. Sikora).
  • Type-depository: MHNG.
  • Combination in C. (Myrmoturba): Santschi, 1917b: 292;
  • combination in C. (Tanaemyrmex): Emery, 1925b: 88.
  • As unavailable (infrasubspecific) name: Santschi, 1911e: 131 (caption of fig. 3d).
  • Subspecies of maculatus: Santschi, 1917b: 292; Santschi, 1919b: 239; Emery, 1920c: 5; Santschi, 1920g: 180; Arnold, 1922: 626; Wheeler, W.M. 1922a: 958, 1042; Emery, 1925b: 88; Santschi, 1925h: 166; Arnold, 1926: 292; Santschi, 1930a: 73; Santschi, 1930b: 76; Santschi, 1935a: 279; Santschi, 1937d: 240.
  • Junior synonym of maculatus: Baroni Urbani, 1972: 126; Bolton, 1995b: 119.
• sarmentus. Camponotus (Myrmoturba) maculatus subsp. sarmentus Emery, 1920c: 14, fig. 2 (s.w.) DEMOCRATIC REPUBLIC OF CONGO.
  • Type-material: syntype major and minor workers (numbers not stated).
  • Type-locality: Democratic Republic of Congo: Ruwenzori, Karewia (F. Stuhlmann).
  • Type-depository: MSNG (perhaps also MNHU).
  • Combination in C. (Tanaemyrmex): Emery, 1925b: 88.
  • Subspecies of maculatus: Emery, 1925b: 88.
  • Junior synonym of maculatus: Baroni Urbani, 1972: 126; Bolton, 1995b: 122.
• schereri. Camponotus maculatus subsp. schereri Forel, 1911e: 289 (s.w.q.) LIBERIA.
  • Type-material: syntype major and minor workers, syntype queen(s) (numbers not stated).
  • Type-locality: Liberia: Nabena (Scherer).
  • Type-depository: ZSBS (perhaps also MHNG).
  • Combination in C. (Myrmoturba): Santschi, 1915c: 280;
  • combination in C. (Tanaemyrmex): Emery, 1925b: 88.
  • Subspecies of maculatus: Santschi, 1914d: 379; Santschi, 1915c: 280; Emery, 1920c: 5; Wheeler, W.M. 1922a: 959; Emery, 1925b: 88.
  • Junior synonym of maculatus: Baroni Urbani, 1972: 126; Bolton, 1995b: 122.
• schultzei. Camponotus maculatus var. schultzei Forel, 1912j: 179 (s.w.m.) UGANDA.
  • Type-material: syntype major and minor workers, syntype male(s) (numbers not stated).
  • Type-locality: Uganda: Entebbe, iii.1912 (H. Schultze).
  • Type-depository: MHNG.
  • Menozzi, 1933a: 110 (q.).
  • As unavailable (infrasubspecific) name: Santschi, 1915c: 279; Emery, 1920c: 5; Wheeler, W.M. 1922a: 957.
  • Subspecies of maculatus: Santschi, 1914b: 130; Menozzi, 1933a: 110; Menozzi, 1939c: 106.
  • Junior synonym of melanocnemis: Forel, 1914d: 250; Emery, 1925b: 87.
  • Junior synonym of maculatus: Baroni Urbani, 1972: 124; Bolton, 1995b: 110.
• semispicatus. Camponotus (Tanaemyrmex) maculatus var. semispicatus Emery, 1925b: 87.
  • [First available use of Camponotus (Myrmoturba) maculatus st. mathildae var. semispicata Santschi, 1914b: 130 (s.w.q.m.) KENYA; unavailable (infrasubspecific) name.]
  • Type-material: syntype major and minor workers, syntype queens, syntype males (numbers not stated).
  • Type-localities: Kenya (“Afrique orientale anglaise”): Nairobi, Kikuyu land, 1700 m., st. no. 11, xi. 1911 (Ch. Alluaud & R. Jeannel) (invalid restriction of type-locality by Wheeler, W.M. 1922a: 957; no lectotype designated), Kenya: Naivasha, 1900 m., st. no. 14, xii.1911 (Ch. Alluaud & R. Jeannel), Kenya: Gilgil, 1980 m., st. no. 15, xii.1911 (Ch. Alluaud & R. Jeannel), Kenya: Nakuru, 1280 m., st. no. 17, xii.1911 (Ch. Alluaud & R. Jeannel), Kenya: Kijabe, Kikuyu escarpment, 2100 m., st. no. 27, xii.1911 (Ch. Alluaud & R. Jeannel), Kenya: nr Nairobi, Masai steppe, 1904 (no collector’s name, probably Ch. Alluaud).
  • Type-depository: NHMB.
  • Menozzi, 1930b: 116 (q.).
  • As unavailable (infrasubspecific) name: Santschi, 1915c: 279; Emery, 1920c: 5; Wheeler, W.M. 1922a: 957.
  • Subspecies of maculatus: Emery, 1925b: 87; Menozzi, 1930b: 115; Menozzi, 1939c: 107; Menozzi & Consani, 1952: 69; Bernard, 1953b: 261.
  • Junior synonym of maculatus: Baroni Urbani, 1972: 124; Bolton, 1995b: 123.
• sudanicus. Camponotus (Myrmoturba) maculatus subsp. sudanicus Weber, 1943c: 385 (s.w.) SOUTH SUDAN.
  • Type-material: syntype major and minor workers (numbers not stated, “one colony”).
  • Type-locality: South Sudan (“Anglo-Egyptian Sudan”): Imatong Mts, 6300 ft, 2.viii.1939, no. 1396 (N.A. Weber).
  • Type-depository: MCZC.
  • Junior synonym of maculatus: Baroni Urbani, 1972: 126; Bolton, 1995b: 125.
• thomensis. Camponotus (Myrmoturba) maculatus var. thomensis Santschi, 1920i: 3 (s.w.m.) SÃO TOMÉ & PRÍNCIPE (São Tomé).
  • Type-material: syntype major and minor workers, syntype male(s) (numbers not stated).
  • Type-locality: São Tomé & Principe: São Tomé I., no. 701 (A.F. de Seabra).
  • Type-depository: NHMB.
  • [Misspelled as thomeensis by Santschi, 1939f: 167.]
  • Subspecies of maculatus: Wheeler, W.M. 1922a: 951; Emery, 1925b: 87; Santschi, 1939f: 167.
  • Junior synonym of maculatus: Baroni Urbani, 1972: 125; Bolton, 1995b: 127.
• tuckeri. Camponotus (Myrmoturba) maculatus var. tuckeri Santschi, 1932a: 391 (s.w.m.) NAMIBIA.
  • Type-material: syntype major and minor workers, syntype males (numbers not stated).
  • Type-locality: Namibia (“SW Africa”): Aanaus, 18.xii.1913 (no collector’s name).
  • Type-depository: SAMC (perhaps also NHMB).
  • Junior synonym of maculatus: Baroni Urbani, 1972: 125; Bolton, 1995b: 128.
• zumpti. Camponotus (Myrmoturba) maculatus var. zumpti Santschi, 1937b: 103 (s.w.) CAMEROON.
  • Type-material: 4 syntype major workers, 2 syntype minor workers.
  • Type-locality: Cameroon: Missellele, viii-xi.1935 (F. Zumpt).
  • Type-depository: NHMB.
  • Junior synonym of maculatus: Baroni Urbani, 1972: 125; Bolton, 1995b: 130.

Taxonomic Notes

Taylor et al. (2018): Camponotus lohieri Santschi, 1913, new status (here, originally described as Camponotus (Myrmoturba) maculatus Fab. stirps melanocnemis Sants. v. lohieri n. var. Santschi, 1913; raised status to Camponotus maculatus F., var. lohieri Sant., Emery, 1915g: 22), CASENT0905268), Haut-Dahomey, DESANTI.

This treatment of C. lohieri as a valid species by Taylor et al. (2018) was proposed without comment or justification, has has not been accepted by subsequent authors.

Taylor et al. (2018) treat Camponotus proletaria as a full, distinct species but provide no justification for the change. Subsequent authors have not followed this treatment.

References

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